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Male
Greater
Prairie-Chicken
in mating display |
Greater Prairie-Chicken
(Tympanuchus cupido) North
American distribution of Pinnated Grouse (Greater and Lesser Prairie-Chickens)
once ranged from the Gulf Coast to Southern Canada, and from the Atlantic
Seaboard to the Rocky Mountains, but this range has been steadily decreasing
over the centuries (see attached Map 1 from Johnsguard, 1973). Of the three
subspecies of Greater Prairie-Chickens, the Heath Hen is extinct, and the
Attwater's Prairie-Chicken on the TX coast is hovering on the brink of extinction.
The bulk of Greater Prairie-Chickens are today found in tall grass and mixed
grass prairie regions of KS, NE, and SD with fringe and remnant populations
extending into other states and provinces such as OK, MO, ND, and Ontario.
In other states such as WI, IL, MI, and MN, populations continue to exist
through reintroduction efforts. On the fringe of the distribution such as
in OK, numbers have gradually decreased to a low in the mid 90's, but have
shown gradual increases during the last three years. While Greater Prairie-Chicken
numbers appear stable in some states comprising the heart of the distribution
region, the latest counts indicate that today there are only about one-third
as many of this species as there were just 30 years ago. (Source: Ron Westermeier
and Sharon Gough, Illinois Natural History Survey and Missouri Department
of Conservation respectively, 1999 Report on the National Outlook and Conservation
Needs for Greater Prairie-Chickens). |
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| Attwater Prairie-Chicken
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GPC
with tracking collar |
Extinct
Heath Hen |
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Habitat

Range

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Links of Interest
The Attwater Prairie-Chicken
USGS
Patuxent Wildlife Research Center
Sutton
Avian Research Center
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Greater Sage-Grouse
www.joelsartore.com |
Sage Grouse
(Centrocercus urophasianus)
General Description and Species Variation: The Greater Sage-Grouse
(Centrocercus urophasianus) is the largest grouse in North America.
Its very large size (70-75 cm long, males = 2.7-3.3 kg, females = 1.4-1.9
kg), long pointed tail, and distinctive drab gray and white plumage distinguish
the Greater sage-grouse from all other North American grouse, except the
Gunnison sage-grouse (Centrocercus minimus). Recently recognized
as a distinct species, the Gunnison sage-grouse is similar in appearance
but about 30% smaller (males = 1.6-2.4 kg, females = 1.0-1.4 kg) and lighter
in color than the Greater sage-grouse. In addition, male Gunnison sage-grouse
have shorter, more distinctively barred tail feathers than male Greater
sage-grouse. Males
of both species are larger than the females and possess prominent white
upper breast feathers. During the breeding season, males develop discernible
air sacs on the breast and specialized ornamental contour feathers called
filoplumes that arise from the dorsal base of the neck. The filoplumes
of male Gunnison sage-grouse are longer and thicker than those of Greater
sage-grouse males. There are 2 weakly differentiated subspecies of Greater
sage-grouse.
Diet: Sagebrush leaves dominate the diet of adult sage grouse throughout
the year, but are especially important during winter. Forbs are consumed
by hens during prelaying and by all age and sex classes during summer.
Insects are critical for juveniles during the first 3-4 weeks of life,
with forbs increasing in the diet as the juveniles age. Insects also are
utilized by adults during spring and summer.
Breeding and Nesting Characteristics: Males gather on traditional
breeding areas known as leks or strutting grounds where they perform elaborate
displays and vocalizations to attract females for breeding. The strut
display differs between the 2 species. Male Gunnison sage-grouse display
at a slower rate, pop their air sacs more frequently, toss their heads
more vigorously to display their filoplumes, and terminate the display
with a tail wag. Yearling males seldom attempt to breed and only 10-15%
of the adult males actually are successful at mating on the lek. Females
may nest anywhere from 2-20 km from the lek where they were bred. They
are ground nesters and prefer to construct their nest under sagebrush
plants with higher residual cover than occurs randomly within the sagebrush
community. Females lay an average of 7-9 eggs that are olive-buff to greenish-brown
with small dark brown spots. The incubation period lasts 25-29 days. Nesting
success is highly variable, but usually averages between 40-60%. Both
adult and subadult hens will attempt to nest, but adults tend to be more
successful than subadults and exhibit a greater tendency to renest if
the first nest is destroyed. On average, 40-50% of the hens will successfully
hatch a clutch of eggs. Sage grouse raise one brood per year. The chicks
are fully feathered and capable of foraging on their own at hatching.
They can sustain short flights at 10-14 days of age and become independent
of the brood hen at 10-12 weeks of age.
Predation:
Although sage-grouse congregate on leks and nest on the ground,
behaviors that should subject them to higher predation rates, their
larger size limits the number of predators (especially avian predators)
that prey upon them. Thus, sage grouse tend to have higher survival
rates than other species of grouse with females (55-75%) surviving
at a higher rate than males (35-60%).
Current Problems and Threats:
The distribution and abundance of sage-grouse have declined dramatically
throughout the range for both species. The Gunnison sage-grouse is
currently classified as a "candidate species" and petitions
have been filed to list the Greater sage-grouse. The primary threats
to sage grouse population include habitat loss, degradation, and fragmentation
due to conversion of native sagebrush communities to croplands and
pasture for livestock, excessive livestock grazing within remaining
sagebrush communities, herbicide treatment, invasion of non-native
plants, encroachment of conifers, unnatural fire regimes, and energy
development. |
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Habitat
Habitat: As its name implies, sage grouse are
closely associated with the sagebrush ecosystem of western North America.
There is tremendous natural variation in sagebrush communities within
this ecosystem to which the sage grouse have adapted.
Range
Range: The current range of
the Greater sage-grouse includes extreme southeast Alberta and southwest
Saskatchewan, extreme southwest North Dakota and northwest South Dakota,
much of Montana and Wyoming, western Colorado, parts of southern and eastern
Idaho, portions of north, northeast and south Utah, northern Nevada, extreme
east to northeast California, southeast Oregon, and north-central Washington.
Gunnison sage-grouse occur in small isolated populations in southwest
Colorado and southeast Utah.

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Links of Interest
USGS
Patuxent Wildlife Research Center
BLM
report on Greater Sage-Grouse and Sagebruse Steppe Ecosystems-- Management
Guidelines
USGS Sagemap Portal
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Lesser
Prairie-Chicken (Tympanuchus pallidicinctus)
| General Description:
Medium-sized grouse weighing 700-800 g. Total length 38-41 cm. Sexes
and plumages similar throughout year. In adults, plumage is uniformly
barred with alternating brown and buffy white bands. Upperparts
are darker and more richly colored than underparts. Chin and throat
largely unmarked. Tail short, rounded and brownish black. Males
display conspicuous yellow eye-combs and dull red gular sacs during
courtship. Males also have elongated pinnae on each side of the
neck, which are held erect during courtship display. Females have
shorter pinnae. Immatures similar to adults, but more richly colored,
especially on throat. Total length, and lengths of wing-chord, pinnae,
and tail, greater in males than females, and within each sex tend
to be greater in adults than in yearlings.
General Food Types: Diet consists of insects,
seeds, leaves, buds, and cultivated grain crops. Juveniles <10
weeks old feed primarily on insects, principally grasshoppers (Orthoptera)
and beetles.
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Daily Routine:
Fall/Winter: Birds assemble into mixed flocks feeding primarily
in shinnery oak-grasslands, but may also feed on waste grains.
Spring/Summer: During breeding season, males congregate on
traditional lek sites to attract females to mate. Nests are initiated
mid-April through late May, typically within 2 weeks of lek attendance
and copulation. Hatching peaks late May-mid June throughout range.
Re-nests (if initial clutch is lost) initiated mid-May-early June,
with hatching mid-June-early July. Estimates of average clutch size
range from 10-12 eggs. Eggs typically ovate, averaging 42.01 (range
40.40-43.93) x 31.60 mm (range 29.17-33.15, n = 86). Egg color varies
from cream to ivory yellow, sometimes dotted with pale brown or olive
spots. Chicks are precocial and nidifugous, leaving nest with hen
within 24 hours of hatching. Chicks capable of short flights at 2
weeks of age. Loss of brood integrity occurs at 12-15 weeks, coinciding
with fall dispersal.
Primary Predators: Predators of adults and chicks include
Rough-legged Hawk (Buteo lagopus), Red-tailed Hawk (Buteo jamaicensis),
Prairie Falcon (Falco mexicanus), Cooper's Hawk (Accipiter cooperii),
Northern Harrier (Circus cyaneus), Ferruginous Hawk (Buteo regalis),
Golden Eagle (Aquila chrysaetos), Great Horned Owl (Bubo virginianus),
coyote (Canis latrans), and badger (Taxidea taxus). Nest predators
include Chihuahuan Raven (Corvus cryptoleucus), coyote, badger,
striped skunk (Mephitis mephitis), ground squirrel (Spermophilus
spilosoma), and bull snake (Pituophis melanoleucus).
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| Primary Problems:
Population declines since 1800s have resulted from conversion, degradation,
and fragmentation of habitat through extrinsic factors, including
drought, conversion of rangeland to cropland, overgrazing by domestic
livestock, chemical control of sand sagebrush and shinnery oak on
rangelands, and oil and gas development.
Literature
cited. |
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Habitat:
Inhabits short- to mid-grass prairies interspersed with shinnery
oak (Quercus havardii) or sand sagebrush (Artemisia filifolia).

Range:
Extends from western Kansas and southeastern Colorado, south to
the northwestern Oklahoma and northern Texas panhandles and eastern
New Mexico.

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Dancing
Columbia Sharp-tailed Grouse |
Sharp-tailed
Grouse (Tympanuchus phasianellus)
| General Description:
Sharp-tailed Grouse have a mottled, light brown appearance. Distinguishing
features include a short, pointed tail, white spots on the wings,
and dark V-shaped markings on the breast feathers. There are 6 subspecies
of sharp-tailed grouse in North America ranging from the Great Lakes
states west to Alaska and south to Colorado. Sharp-tailed grouse
are most closely related to, but smaller than, the greater and lesser
prairie-chicken. Adult sharptails average 42 to 48 cm in length
and weigh anywhere from 596 to 1,031 g depending on subspecies,
age, sex, and season of the year. Males weigh more than females,
and within sexes, adults weigh more than subadults. Unless the males
are displaying and exposing their pinkish to violet-colored air
sacs, they appear similar in size, shape, and color to females.
Birds
in hand can be sexed by the presence (females) or absence (males)
of traverse barring on the central tail feathers and crown feathers.
General Food Types: In all habitats, sharptails
consume a variety of forbs, fruits, seeds, grains, insects, and
buds (primarily during winter).
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| Breeding and Nesting Characteristics:
During spring, males gather on traditional breeding areas called
leks or dancing grounds where they perform elaborate courtship displays
and vocalizations to attract females. Leks are typically located
on knolls, benches, or ridge tops that are slightly higher in elevation
and support lower growing vegetation than the surrounding terrain.
Leks may be used for many years, even decades, and form the hub
of all activities associated with breeding. Most females will nest
and raise their broods within 2 km of the lek where they were bred.
The female constructs a well-concealed, rudimentary nest bowl on
the ground that is lined with dried vegetation and body feathers.
Females lay an average of 10-12 eggs that are light to dark brown
with well-defined reddish-brown spots. The incubation period is
about 24 days. Nesting success (% of hens that hatch 1 egg) ranges
from 40-70%. Both adult and subadult hens attempt to nest, but adults
tend to be more successful than subadults and exhibit a greater
tendency to renest if the first nest is destroyed. Sharp-tailed
grouse raise one brood per year. The chicks are precocial at hatching,
which means they are fully feathered and capable of foraging on
their own. They can fly at about 10 days of age and become independent
of the brood hen between 10 and 12 weeks of age.
Primary Problems: Because sharptails gather
on leks and nest on the ground, they are exceptionally vulnerable
to predation. Annual mortality rates frequently exceed 50%. Populations
in Canada (except British Columbia), Alaska, North Dakota, South
Dakota, Nebraska, and Montana are more secure than populations in
the Great Lakes and intermountain regions of the United States.
The primary threats to sharp-tailed grouse populations include the
loss and degradation of habitats due to conversion of native shrub
and grasslands to croplands, excessive grazing by livestock, herbicide
treatments, fire suppression, invasion of non-native plants, removal
of trees and shrubs in riparian areas, invasion of conifers, and
urban development.
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| Habitat:
Preferred habitat varies from mixed grass prairies to sagebrush
to post-burn, aspen forest grasslands. Historically, this species
was found in shrub-steppe, grassland, and mixed shrub habitats,
but more recently populations have adapted to using croplands, Conservation
Reserve Program lands, and mine reclamation lands to varying degrees.
Habitat requirements are most restricted during winter when the
species is dependent on riparian areas, deciduous shrub thickets,
and deciduous woodlands. Sharptails use flat to rolling terrain
comprised of open shrub-dominated habitats and grasslands for breeding,
nesting, and brood-rearing.

Range:
In one of its six subspecific forms, plains, Columbian, prairie,
northern, Alaskan, and an undescribed race south of Yellow Knife,
this grouse (see attached Map 3 from Johnsguard, 1973) has had the
broadest distribution of any grouse species in North America. It
is found in the northern U.S. and Canada primarily in British Columbia,
Alberta, Saskatchewan, Manitoba, Ontario, Yukon Territory, AK, MT,
WY, ND, SD, NE, MN, WI, and WA with fringes or pockets also in CO,
UT, ID, OR, NM, KS, and Quebec. Throughout the area in Alaska, Canada,
and the mid-west, Sharp-tailed Grouse remain fairly secure. Like
the range of Prairie-Chickens, however, overall the range of Sharp-tailed
Grouse has shrunk considerably during the last century. In particular,
the western or Columbian subspecies and the eastern or prairie subspecies
have suffered severe declines in numbers. The Columbian Sharp-tailed
Grouse, was the subject of a 1995 petition to the USFWS by the Biological
Diversity Legal Foundation (Boulder, CO) for consideration on the
THREATENED and endangered species list.

Links of Interest
USGS
Patuxent Wildlife Research Center
USGS
Northern Prairie Wildlife Research Center
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