Male Greater
Prairie-Chicken
in mating display
Greater Prairie-Chicken (Tympanuchus cupido)

North American distribution of Pinnated Grouse (Greater and Lesser Prairie-Chickens) once ranged from the Gulf Coast to Southern Canada, and from the Atlantic Seaboard to the Rocky Mountains, but this range has been steadily decreasing over the centuries (see attached Map 1 from Johnsguard, 1973). Of the three subspecies of Greater Prairie-Chickens, the Heath Hen is extinct, and the Attwater's Prairie-Chicken on the TX coast is hovering on the brink of extinction. The bulk of Greater Prairie-Chickens are today found in tall grass and mixed grass prairie regions of KS, NE, and SD with fringe and remnant populations extending into other states and provinces such as OK, MO, ND, and Ontario. In other states such as WI, IL, MI, and MN, populations continue to exist through reintroduction efforts. On the fringe of the distribution such as in OK, numbers have gradually decreased to a low in the mid 90's, but have shown gradual increases during the last three years. While Greater Prairie-Chicken numbers appear stable in some states comprising the heart of the distribution region, the latest counts indicate that today there are only about one-third as many of this species as there were just 30 years ago. (Source: Ron Westermeier and Sharon Gough, Illinois Natural History Survey and Missouri Department of Conservation respectively, 1999 Report on the National Outlook and Conservation Needs for Greater Prairie-Chickens).
     
   
Attwater Prairie-Chicken GPC with tracking collar Extinct Heath Hen
   


Habitat

Range


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Links of Interest

The Attwater Prairie-Chicken

USGS Patuxent Wildlife Research Center

Sutton Avian Research Center

 


   
 
Greater Sage-Grouse www.joelsartore.com

Sage Grouse (Centrocercus urophasianus)

General Description and Species Variation: The Greater Sage-Grouse (Centrocercus urophasianus) is the largest grouse in North America. Its very large size (70-75 cm long, males = 2.7-3.3 kg, females = 1.4-1.9 kg), long pointed tail, and distinctive drab gray and white plumage distinguish the Greater sage-grouse from all other North American grouse, except the Gunnison sage-grouse (Centrocercus minimus). Recently recognized as a distinct species, the Gunnison sage-grouse is similar in appearance but about 30% smaller (males = 1.6-2.4 kg, females = 1.0-1.4 kg) and lighter in color than the Greater sage-grouse. In addition, male Gunnison sage-grouse have shorter, more distinctively barred tail feathers than male Greater sage-grouse. Males of both species are larger than the females and possess prominent white upper breast feathers. During the breeding season, males develop discernible air sacs on the breast and specialized ornamental contour feathers called filoplumes that arise from the dorsal base of the neck. The filoplumes of male Gunnison sage-grouse are longer and thicker than those of Greater sage-grouse males. There are 2 weakly differentiated subspecies of Greater sage-grouse.

Diet: Sagebrush leaves dominate the diet of adult sage grouse throughout the year, but are especially important during winter. Forbs are consumed by hens during prelaying and by all age and sex classes during summer. Insects are critical for juveniles during the first 3-4 weeks of life, with forbs increasing in the diet as the juveniles age. Insects also are utilized by adults during spring and summer.

Breeding and Nesting Characteristics: Males gather on traditional breeding areas known as leks or strutting grounds where they perform elaborate displays and vocalizations to attract females for breeding. The strut display differs between the 2 species. Male Gunnison sage-grouse display at a slower rate, pop their air sacs more frequently, toss their heads more vigorously to display their filoplumes, and terminate the display with a tail wag. Yearling males seldom attempt to breed and only 10-15% of the adult males actually are successful at mating on the lek. Females may nest anywhere from 2-20 km from the lek where they were bred. They are ground nesters and prefer to construct their nest under sagebrush plants with higher residual cover than occurs randomly within the sagebrush community. Females lay an average of 7-9 eggs that are olive-buff to greenish-brown with small dark brown spots. The incubation period lasts 25-29 days. Nesting success is highly variable, but usually averages between 40-60%. Both adult and subadult hens will attempt to nest, but adults tend to be more successful than subadults and exhibit a greater tendency to renest if the first nest is destroyed. On average, 40-50% of the hens will successfully hatch a clutch of eggs. Sage grouse raise one brood per year. The chicks are fully feathered and capable of foraging on their own at hatching. They can sustain short flights at 10-14 days of age and become independent of the brood hen at 10-12 weeks of age.

Predation: Although sage-grouse congregate on leks and nest on the ground, behaviors that should subject them to higher predation rates, their larger size limits the number of predators (especially avian predators) that prey upon them. Thus, sage grouse tend to have higher survival rates than other species of grouse with females (55-75%) surviving at a higher rate than males (35-60%).

Current Problems and Threats: The distribution and abundance of sage-grouse have declined dramatically throughout the range for both species. The Gunnison sage-grouse is currently classified as a "candidate species" and petitions have been filed to list the Greater sage-grouse. The primary threats to sage grouse population include habitat loss, degradation, and fragmentation due to conversion of native sagebrush communities to croplands and pasture for livestock, excessive livestock grazing within remaining sagebrush communities, herbicide treatment, invasion of non-native plants, encroachment of conifers, unnatural fire regimes, and energy development.
   


Habitat

Habitat:
As its name implies, sage grouse are closely associated with the sagebrush ecosystem of western North America. There is tremendous natural variation in sagebrush communities within this ecosystem to which the sage grouse have adapted.

Range

Range: The current range of the Greater sage-grouse includes extreme southeast Alberta and southwest Saskatchewan, extreme southwest North Dakota and northwest South Dakota, much of Montana and Wyoming, western Colorado, parts of southern and eastern Idaho, portions of north, northeast and south Utah, northern Nevada, extreme east to northeast California, southeast Oregon, and north-central Washington. Gunnison sage-grouse occur in small isolated populations in southwest Colorado and southeast Utah.


click to enlarge

Links of Interest

USGS Patuxent Wildlife Research Center

BLM report on Greater Sage-Grouse and Sagebruse Steppe Ecosystems-- Management Guidelines

USGS Sagemap Portal




 

Lesser Prairie-Chicken (Tympanuchus pallidicinctus)

General Description: Medium-sized grouse weighing 700-800 g. Total length 38-41 cm. Sexes and plumages similar throughout year. In adults, plumage is uniformly barred with alternating brown and buffy white bands. Upperparts are darker and more richly colored than underparts. Chin and throat largely unmarked. Tail short, rounded and brownish black. Males display conspicuous yellow eye-combs and dull red gular sacs during courtship. Males also have elongated pinnae on each side of the neck, which are held erect during courtship display. Females have shorter pinnae. Immatures similar to adults, but more richly colored, especially on throat. Total length, and lengths of wing-chord, pinnae, and tail, greater in males than females, and within each sex tend to be greater in adults than in yearlings.

General Food Types: Diet consists of insects, seeds, leaves, buds, and cultivated grain crops. Juveniles <10 weeks old feed primarily on insects, principally grasshoppers (Orthoptera) and beetles.

Daily Routine:
Fall/Winter
: Birds assemble into mixed flocks feeding primarily in shinnery oak-grasslands, but may also feed on waste grains.
Spring/Summer: During breeding season, males congregate on traditional lek sites to attract females to mate. Nests are initiated mid-April through late May, typically within 2 weeks of lek attendance and copulation. Hatching peaks late May-mid June throughout range. Re-nests (if initial clutch is lost) initiated mid-May-early June, with hatching mid-June-early July. Estimates of average clutch size range from 10-12 eggs. Eggs typically ovate, averaging 42.01 (range 40.40-43.93) x 31.60 mm (range 29.17-33.15, n = 86). Egg color varies from cream to ivory yellow, sometimes dotted with pale brown or olive spots. Chicks are precocial and nidifugous, leaving nest with hen within 24 hours of hatching. Chicks capable of short flights at 2 weeks of age. Loss of brood integrity occurs at 12-15 weeks, coinciding with fall dispersal.

Primary Predators: Predators of adults and chicks include Rough-legged Hawk (Buteo lagopus), Red-tailed Hawk (Buteo jamaicensis), Prairie Falcon (Falco mexicanus), Cooper's Hawk (Accipiter cooperii), Northern Harrier (Circus cyaneus), Ferruginous Hawk (Buteo regalis), Golden Eagle (Aquila chrysaetos), Great Horned Owl (Bubo virginianus), coyote (Canis latrans), and badger (Taxidea taxus). Nest predators include Chihuahuan Raven (Corvus cryptoleucus), coyote, badger, striped skunk (Mephitis mephitis), ground squirrel (Spermophilus spilosoma), and bull snake (Pituophis melanoleucus).

Primary Problems: Population declines since 1800s have resulted from conversion, degradation, and fragmentation of habitat through extrinsic factors, including drought, conversion of rangeland to cropland, overgrazing by domestic livestock, chemical control of sand sagebrush and shinnery oak on rangelands, and oil and gas development.

Literature cited.


Habitat:


Inhabits short- to mid-grass prairies interspersed with shinnery oak (Quercus havardii) or sand sagebrush (Artemisia filifolia).

Range:

Extends from western Kansas and southeastern Colorado, south to the northwestern Oklahoma and northern Texas panhandles and eastern New Mexico.


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Dancing Columbia Sharp-tailed Grouse

Sharp-tailed Grouse (Tympanuchus phasianellus)

General Description: Sharp-tailed Grouse have a mottled, light brown appearance. Distinguishing features include a short, pointed tail, white spots on the wings, and dark V-shaped markings on the breast feathers. There are 6 subspecies of sharp-tailed grouse in North America ranging from the Great Lakes states west to Alaska and south to Colorado. Sharp-tailed grouse are most closely related to, but smaller than, the greater and lesser prairie-chicken. Adult sharptails average 42 to 48 cm in length and weigh anywhere from 596 to 1,031 g depending on subspecies, age, sex, and season of the year. Males weigh more than females, and within sexes, adults weigh more than subadults. Unless the males are displaying and exposing their pinkish to violet-colored air sacs, they appear similar in size, shape, and color to females. Birds in hand can be sexed by the presence (females) or absence (males) of traverse barring on the central tail feathers and crown feathers.

General Food Types: In all habitats, sharptails consume a variety of forbs, fruits, seeds, grains, insects, and buds (primarily during winter).

Breeding and Nesting Characteristics: During spring, males gather on traditional breeding areas called leks or dancing grounds where they perform elaborate courtship displays and vocalizations to attract females. Leks are typically located on knolls, benches, or ridge tops that are slightly higher in elevation and support lower growing vegetation than the surrounding terrain. Leks may be used for many years, even decades, and form the hub of all activities associated with breeding. Most females will nest and raise their broods within 2 km of the lek where they were bred. The female constructs a well-concealed, rudimentary nest bowl on the ground that is lined with dried vegetation and body feathers. Females lay an average of 10-12 eggs that are light to dark brown with well-defined reddish-brown spots. The incubation period is about 24 days. Nesting success (% of hens that hatch 1 egg) ranges from 40-70%. Both adult and subadult hens attempt to nest, but adults tend to be more successful than subadults and exhibit a greater tendency to renest if the first nest is destroyed. Sharp-tailed grouse raise one brood per year. The chicks are precocial at hatching, which means they are fully feathered and capable of foraging on their own. They can fly at about 10 days of age and become independent of the brood hen between 10 and 12 weeks of age.

Primary Problems: Because sharptails gather on leks and nest on the ground, they are exceptionally vulnerable to predation. Annual mortality rates frequently exceed 50%. Populations in Canada (except British Columbia), Alaska, North Dakota, South Dakota, Nebraska, and Montana are more secure than populations in the Great Lakes and intermountain regions of the United States. The primary threats to sharp-tailed grouse populations include the loss and degradation of habitats due to conversion of native shrub and grasslands to croplands, excessive grazing by livestock, herbicide treatments, fire suppression, invasion of non-native plants, removal of trees and shrubs in riparian areas, invasion of conifers, and urban development.

Habitat:

Preferred habitat varies from mixed grass prairies to sagebrush to post-burn, aspen forest grasslands. Historically, this species was found in shrub-steppe, grassland, and mixed shrub habitats, but more recently populations have adapted to using croplands, Conservation Reserve Program lands, and mine reclamation lands to varying degrees. Habitat requirements are most restricted during winter when the species is dependent on riparian areas, deciduous shrub thickets, and deciduous woodlands. Sharptails use flat to rolling terrain comprised of open shrub-dominated habitats and grasslands for breeding, nesting, and brood-rearing.

Range:

In one of its six subspecific forms, plains, Columbian, prairie, northern, Alaskan, and an undescribed race south of Yellow Knife, this grouse (see attached Map 3 from Johnsguard, 1973) has had the broadest distribution of any grouse species in North America. It is found in the northern U.S. and Canada primarily in British Columbia, Alberta, Saskatchewan, Manitoba, Ontario, Yukon Territory, AK, MT, WY, ND, SD, NE, MN, WI, and WA with fringes or pockets also in CO, UT, ID, OR, NM, KS, and Quebec. Throughout the area in Alaska, Canada, and the mid-west, Sharp-tailed Grouse remain fairly secure. Like the range of Prairie-Chickens, however, overall the range of Sharp-tailed Grouse has shrunk considerably during the last century. In particular, the western or Columbian subspecies and the eastern or prairie subspecies have suffered severe declines in numbers. The Columbian Sharp-tailed Grouse, was the subject of a 1995 petition to the USFWS by the Biological Diversity Legal Foundation (Boulder, CO) for consideration on the THREATENED and endangered species list.

Links of Interest

USGS Patuxent Wildlife Research Center

USGS Northern Prairie Wildlife Research Center

 

 

     
     
     
 
 

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